第6章

IntheCalamariesandthePoulpstheink-bagisplacedintheupperpartofthebody,incloseproximitytothemytis,whereasintheSepiaitislowerdown,againstthestomach。FortheSepiahasamoreplentifulsupplyofinkthantherest,inasmuchasitmakesmoreuseofit。Thereasonsforthisare,firstly,thatitlivesneartheshore,and,secondly,thatithasnoothermeansofprotection;whereasthePoulphasitslongtwiningfeettouseinitsdefence,andis,moreover,endowedwiththepowerofchangingcolour。Thischangingofcolour,likethedischargeofink,occursastheresultoffright。AstotheCalamary,itlivesfaroutatsea,beingtheonlyoneoftheCephalopodathatdoesso;andthisgivesitprotection。ThesethenarethereasonswhytheinkismoreabundantintheSepiathanintheCalamary,andthisgreaterabundanceexplainsthelowerposition;foritallowstheinktobeejectedwitheaseevenfromadistance。Theinkitselfisofanearthycharacter,inthisresemblingthewhitedepositonthesurfaceofabird’sexcrementandtheexplanationinbothcasesisthesame,namely,theabsenceofaurinarybladder。For,indefaultofthis,itistheinkthatservesfortheexcretionoftheearthiestmatter。

AndthisismoreespeciallythecaseintheSepia,becausethereisagreaterproportionofearthinitscompositionthaninthatoftheotherCephalopoda。Theearthycharacterofitsboneisaclearindicationofthis。ForinthePoulpthereisnoboneatall,andintheCalamaryitisthinandcartilaginous。WhythisboneshouldbepresentinsomeCephalopoda,andwantinginothers,andhowitscharactervariesinthosethathaveit,hasnowbeensetforth。

Theseanimals,havingnoblood,areinconsequencecoldandofatimidcharacter。Now,insomeanimals,fearcausesadisturbanceofthebowels,and,inothers,aflowofurinefromthebladder。

Similarlyintheseitproducesadischargeofink,and,thoughtheejectionofthisinkinfright,likethatoftheurine,istheresultofnecessity,and,thoughitisofexcrementalcharacter,yetitisusedbynatureforapurpose,namely,theprotectionandsafetyoftheanimalthatexcretesit。

TheCrustaceaalso,boththeCaraboidformsandtheCrabs,areprovidedwithteeth,namelytheirtwoanteriorteeth;andbetweenthesetheyalsopresentthetongue-likepieceofflesh,ashasindeedbeenalreadymentioned。Directlyaftertheirmouthcomesagullet,which,ifwecomparerelativesizes,isbutsmallinproportiontothebody:andthenastomach,whichintheCarabiandsomeoftheCrabsisfurnishedwithasecondsetofteeth,theanteriorteethbeinginsufficientforadequatemastication。Fromthestomachauniformgutrunsinadirectlinetotheexcrementalvent。

ThepartsdescribedaretobefoundalsoinallthevariousTestacea。Thedegreeofdistinctness,however,withwhichtheyareformedvariesinthedifferentkinds,andthelargerthesizeoftheanimalthemoreeasilydistinguishableareallthesepartsseverally。IntheSea-snails,forexample,wefindteeth,hardandsharp,asbeforementioned,andbetweenthemtheflesh-likesubstance,justasintheCrustaceaandCephalopoda,andagaintheproboscis,which,ashasbeenstated,issomethingbetweenastingandatongue。Directlyafterthemouthcomesakindofbird-likecrop,thenagullet,succeededbyastomach,inwhichisthemecon,asitisstyled;andcontinuouswiththismeconisanintestine,startingdirectlyfromit。ItisthisresidualsubstancewhichappearsinalltheTestaceatoformthemostpalatablemorsel。PurpurasandWhelks,andallotherTestaceathathaveturbinateshells,instructureresembletheSea-snail。ThegeneraandspeciesofTestaceaareverynumerous。Fortherearethosewithturbinateshells,ofwhichsomehavejustbeenmentioned;and,besidesthese,therearebivalvesandunivalves。Thosewithturbinateshellsmay,indeed,afteracertainfashionbesaidtoresemblebivalves。Fortheyallfromtheirverybirthhaveanoperculumtoprotectthatpartoftheirbodywhichisexposedtoview。ThisisthecasewiththePurpuras,withWhelks,withtheNerites,andthelike。Wereitnotforthis,thepartwhichisundefendedbytheshellwouldbeveryliabletoinjurybycollisionwithexternalobjects。Theunivalvesalsoarenotwithoutprotection。Forontheirdorsalsurfacetheyhaveashell,andbytheundersurfacetheyattachthemselvestotherocks,andsoafteramannerbecomebivalved,therockrepresentingthesecondvalve。OfthesetheanimalsknownasLimpetsareanexample。Thebivalves,scallopsandmussels,forinstance,areprotectedbythepowertheyhaveofclosingtheirvalves;andtheTurbinatabytheoperculumjustmentioned,whichtransformsthem,asitwere,cromunivalvesintobivalves。Butofallthereisnonesoperfectlyprotectedasthesea-urchin。Forherethereisaglobularshellwhichenclosesthebodycompletely,andwhichis,moreover,setwithsharpspines。Thispeculiaritydistinguishesthesea-urchinfromallotherTestacea,ashasalreadybeenmentioned。

ThestructureoftheTestaceaandoftheCrustaceaisexactlythereverseofthatoftheCephalopoda。Forinthelatterthefleshysubstanceisontheoutsideandtheearthysubstancewithin,whereasintheformerthesoftpartsareinsideandthehardpartwithout。

Inthesea-urchin,however,thereisnofleshypartwhatsoever。

AlltheTestaceathen,thosethathavenotbeenmentionedaswellasthosethathave,agreeasstatedinpossessingamouthwiththetongue-likebody,astomach,andaventforexcrement,buttheydifferfromeachotherinthepositionsandproportionsoftheseparts。Thedetails,however,ofthesedifferencesmustbelookedforintheResearchesconcerningAnimalsandthetreatisesonAnatomy。Forwhiletherearesomepointswhichcanbemadeclearbyverbaldescription,thereareotherswhicharemoresuitedforoculardemonstration。

PeculiaramongtheTestaceaarethesea-urchinsandtheanimalsknownasTethya(Ascidians)。Thesea-urchinshavefiveteeth,andinthecentreofthesethefleshybodywhichiscommontoalltheanimalswehavebeendiscussing。Immediatelyafterthiscomesagullet,andthenthestomach,dividedintoanumberofseparatecompartments,whichlooklikesomanydistinctstomachs;forthecavitiesareseparateandallcontainabundantresidualmatter。Theyareall,however,connectedwithoneandthesameoesophagus,andtheyallendinoneandthesameexcrementalvent。Thereisnothingbesidesthestomachofafleshycharacter,ashasalreadybeenstated。Allthatcanbeseenaretheso-calledova,ofwhichthereareseveral,containedeachinaseparatemembrane,andcertainblackbodieswhichhavenoname,andwhich,beginningattheanimal’smouth,arescatteredrounditsbodyhereandtherepromiscuously。Thesesea-urchinsarenotallofonespecies,butthereareseveraldifferentkinds,andinallofthemthepartsmentionedaretobefound。Itisnot,however,ineverykindthattheso-calledovaareedible。Neitherdotheseattaintoanysizeinanyotherspeciesthanthatwithwhichweareallfamiliar。AsimilardistinctionmaybemadegenerallyinthecaseofallTestacea。Forthereisagreatdifferenceintheediblequalitiesofthefleshofdifferentkinds;

andinsome,moreover,theresidualsubstanceknownasthemeconisgoodforfood,whileinothersitisuneatable。Thismeconintheturbinatedgeneraislodgedinthespiralpartoftheshell,whileinunivalves,suchaslimpets,itoccupiesthefundus,andinbivalvesisplacednearthehinge,theso-calledovumlyingontheright;whileontheoppositesideisthevent。Theformerisincorrectlytermedovum,foritmerelycorrespondstowhatinwell-fedsanguineousanimalsisfat;andthusitisthatitmakesitsappearanceinTestaceaatthoseseasonsoftheyearwhentheyareingoodcondition,namely,springandautumn。FornoTestaceacanabideextremesoftemperature,andtheyarethereforeinevilplightinseasonsofgreatcoldorheat。Thisisclearlyshownbywhatoccursinthecaseofthesea-urchins。Forthoughtheovaaretobefoundintheseanimalsevendirectlytheyareborn,yettheyacquireagreatersizethanusualatthetimeoffullmoon;not,assomethink,becausesea-urchinseatmoreatthatseason,butbecausethenightsarethenwarmer,owingtothemoonlight。Forthesecreaturesarebloodless,andsoareunabletostandcoldandrequirewarmth。Thereforeitisthattheyarefoundinbetterconditioninsummerthanatanyotherseason;

andthisallovertheworldexceptinginthePyrrheantidalstrait。

Therethesea-urchinsflourishaswellinwinterasinsummer。Butthereasonforthisisthattheyhaveagreaterabundanceoffoodinthewinter,becausethefishdesertthestraitatthatseason。

Thenumberoftheovaisthesameinallsea-urchins,andisanoddone。Fortherearefiveova,justastherearealsofiveteethandfivestomachs;andtheexplanationofthisistobefoundinthefactthattheso-calledovaarenotreallyova,butmerely,aswassaidbefore,theresultoftheanimal’swell-fedcondition。Oystersalsohaveaso-calledovum,correspondingincharactertothatofthesea-urchins,butexistingonlyononesideoftheirbody。Nowinasmuchasthesea-urchinisofasphericalform,andnotmerelyasinglediskliketheoyster,andinvirtueofitssphericalshapeisthesamefromwhateversideitbeexamined,itsovummustnecessarilybeofacorrespondingsymmetry。Forthesphericalshapehasnottheasymmetryofthedisk-shapedbodyoftheoysters。Forinalltheseanimalstheheadiscentral,butinthesea-urchintheso-calledovumisabove[andsymmetrical,whileintheoysteritisonlyoneside]。Nowthenecessarysymmetrywouldbeobservedweretheovumtoformacontinuousring。Butthismaynotbe。ForitwouldbeinoppositiontowhatprevailsinthewholetribeofTestacea;forinalltheovumisdiscontinuous,andinallexceptingthesea-urchinsasymmetrical,beingplacedonlyononesideofthebody。Owingthentothisnecessarydiscontinuityoftheovum,whichbelongstothesea-urchinasamemberoftheclass,andowingtothesphericalshapeofitsbody,whichisitsindividualpeculiarity,thisanimalcannotpossiblyhaveanevennumberofova。Forweretheyanevennumber,theywouldhavetobearrangedexactlyoppositetoeachother,inpairs,soastokeepthenecessarysymmetry;oneovumofeachpairbeingplacedatoneend,theotherovumattheotherendofatransversediameter。ThisagainwouldviolatetheuniversalprovisioninTestacea。Forbothintheoystersandinthescallopswefindtheovumonlyononesideofthecircumference。Thenumberthenoftheovamustbeuneven,threeforinstance,orfive。Butiftherewereonlythreetheywouldbemuchtoofarapart;while,ifthereweremorethanfive,theywouldcometoformacontinuousmass。Theformerarrangementwouldbedisadvantageoustotheanimal,thelatteranimpossibility。Therecanthereforebeneithermorenorlessthanfive。Forthesamereasonthestomachisdividedintofiveparts,andthereisacorrespondingnumberofteeth。Forseeingthattheovarepresenteachofthemakindofbodyfortheanimal,theirdispositionmustconformtothatofthestomach,seeingthatitisfromthisthattheyderivethematerialfortheirgrowth。Nowiftherewereonlyonestomach,eithertheovawouldbetoofarofffromit,oritwouldbesobigastofillupthewholecavity,andthesea-urchinwouldhavegreatdifficultyinmovingaboutandfindingduenourishmentforitsrepletion。Asthentherearefiveintervalsbetweenthefiveova,soarethereofnecessityfivedivisionsofthestomach,oneforeachinterval。Soalso,andonlikegrounds,therearefiveteeth。Fornatureisthusenabledtoallottoeachstomachalcompartmentandovumitsseparateandsimilartooth。

These,then,arethereasonswhythenumberofovainthesea-urchinisanoddone,andwhythatoddnumberisfive。Insomesea-urchinstheovaareexcessivelysmall,inothersofconsiderablesize,theexplanationbeingthatthelatterareofawarmerconstitution,andsoareabletoconcocttheirfoodmorethoroughly;whileintheformerconcoctionislessperfect,sothatthestomachisfoundfullofresidualmatter,whiletheovaaresmallanduneatable。Thoseofawarmerconstitutionare,moreover,invirtueoftheirwarmthmoregiventomotion,sothattheymakeexpeditionsinsearchoffood,insteadofremainingstationaryliketherest。Asevidenceofthis,itwillbefoundthattheyalwayshavesomethingorotherstickingtotheirspines,asthoughtheymovedmuchabout;fortheyusetheirspinesasfeet。

TheAscidiansdifferbutslightlyfromplants,andyethavemoreofananimalnaturethanthesponges,whicharevirtuallyplantsandnothingmore。Fornaturepassesfromlifelessobjectstoanimalsinsuchunbrokensequence,interposingbetweenthembeingswhichliveandyetarenotanimals,thatscarcelyanydifferenceseemstoexistbetweentwoneighbouringgroupsowingtotheircloseproximity。

Asponge,then,asalreadysaid,intheserespectscompletelyresemblesaplant,thatthroughoutitslifeitisattachedtoarock,andthatwhenseparatedfromthisitdies。Slightlydifferentfromthespongesaretheso-calledHolothuriasandthesea-lungs,asalsosundryothersea-animalsthatresemblethem。Forthesearefreeandunattached。Yettheyhavenofeeling,andtheirlifeissimplythatofaplantseparatedfromtheground。Forevenamongland-plantstherearesomethatareindependentofthesoil,andthatspringupandgrow,eitheruponotherplants,orevenentirelyfree。Such,forexample,istheplantwhichisfoundonParnassus,andwhichsomecalltheEpipetrum。Thisyoumayhanguponapeganditwillyetliveforaconsiderabletime。Sometimesitisamatterofdoubtwhetheragivenorganismshouldbeclassedwithplantsorwithanimals。TheAscidians,forinstance,andthelikesofarresembleplantsasthattheyneverlivefreeandunattached,but,ontheotherhand,inasmuchastheyhaveacertainflesh-likesubstance,theymustbesupposedtopossesssomedegreeofsensibility。

AnAscidianhasabodydividedbyasingleseptumandwithtwoorifices,onewhereittakesinthefluidmatterthatministerstoitsnutrition,theotherwhereitdischargesthesurplusofunusedjuice,forithasnovisibleresidualsubstance,suchashavetheotherTestacea。ThisisitselfaverystrongjustificationforconsideringanAscidian,andanythingelsetheremaybeamonganimalsthatresemblesit,tobeofavegetablecharacter;forplantsalsoneverhaveanyresiduum。AcrossthemiddleofthebodyoftheseAscidiansthererunsathintransversepartition,andhereitisthatwemayreasonablysupposethepartonwhichlifedependstobesituated。

TheAcalephae,orSea-nettles,astheyarevariouslycalled,arenotTestaceaatall,butlieoutsidetherecognizedgroups。Theirconstitution,likethatoftheAscidians,approximatesthemononesidetoplants,ontheothertoanimals。Forseeingthatsomeofthemcandetachthemselvesandcanfastenupontheirfood,andthattheyaresensibleofobjectswhichcomeincontactwiththem,theymustbeconsideredtohaveananimalnature。Thelikeconclusionfollowsfromtheirusingtheasperityoftheirbodiesasaprotectionagainsttheirenemies。But,ontheotherhand,theyarecloselyalliedtoplants,firstlybytheimperfectionoftheirstructure,secondlybytheirbeingabletoattachthemselvestotherocks,whichtheydowithgreatrapidity,andlastlybytheirhavingnovisibleresiduumnotwithstandingthattheypossessamouth。

VerysimilaragaintotheAcalephaearetheStarfishes。Forthesealsofastenontheirprey,andsuckoutitsjuices,andthusdestroyavastnumberofoysters。AtthesametimetheypresentacertainresemblancetosuchoftheanimalswehavedescribedastheCephalopodaandCrustacea,inasmuchastheyarefreeandunattached。

ThesamemayalsobesaidoftheTestacea。

Such,then,isthestructureofthepartsthatministertonutritionandwhicheveryanimalmustpossess。Butbesidestheseorgansitisquiteplainthatineveryanimaltheremustbesomepartorotherwhichshallbeanalogoustowhatinsanguineousanimalsisthepresidingseatofsensation。Whetherananimalhasorhasnotblood,itcannotpossiblybewithoutthis。IntheCephalopodathispartconsistsofafluidsubstancecontainedinamembrane,throughwhichrunsthegulletonitswaytothestomach。Itisattachedtothebodyrathertowardsitsdorsalsurface,andbysomeiscalledthemytis。JustsuchanotherorganisfoundalsointheCrustaceaandtheretooisknownbythesamename。Thispartisatoncefluidandcorporealand,asbeforesaid,istraversedbythegullet。Forhadthegulletbeenplacedbetweenthemytisandthedorsalsurfaceoftheanimal,thehardnessofthebackwouldhaveinterferedwithitsduedilatationintheactofdeglutition。Ontheoutersurfaceofthemytisrunstheintestine;andincontactwiththislatterisplacedtheink-bag,sothatitmayberemovedasfaraspossiblefromthemouthanditsobnoxiousfluidbekeptatadistancefromthenoblerandsovereignpart。Thepositionofthemytisshowsthatitcorrespondstotheheartofsanguineousanimals;foritoccupiestheself-sameplace。Thesameisshownbythesweetnessofitsfluid,whichhasthecharacterofconcoctedmatterandresemblesblood。

IntheTestaceathepresidingseatofsensationisinacorrespondingposition,butislesseasilymadeout。Itshould,however,alwaysbelookedforinsomemidwayposition;namely,insuchTestaceaasarestationary,midwaybetweenthepartbywhichfoodistakeninandthechannelthroughwhicheithertheexcrementorthespermaticfluidisvoided,and,inthosespecieswhicharecapableoflocomotion,invariablymidwaybetweentherightandleftsides。

InInsectsthisorgan,whichistheseatofsensation,lies,aswasstatedinthefirsttreatise,betweentheheadandthecavitywhichcontainsthestomach。Inmostofthemitconsistsofasinglepart;butinothers,forinstanceinsuchashavelongbodiesandresembletheJuli(Millipedes),itismadeupofseveralparts,sothatsuchinsectscontinuetoliveaftertheyhavebeencutinpieces。

Fortheaimofnatureistogivetoeachanimalonlyonesuchdominantpart;andwhensheisunabletocarryoutthisintentionshecausestheparts,thoughpotentiallymany,toworktogetheractuallyasone。Thisismuchmoreclearlymarkedinsomeinsectsthaninothers。

Thepartsconcernedinnutritionarenotalikeinallinsects,butshowconsiderablediversity。Thussomehavewhatiscalledastinginthemouth,whichisakindofcompoundinstrumentthatcombinesinitselfthecharacterofatongueandoflips。Inothersthathavenosuchinstrumentinfrontthereisapartinsidethemouththatanswersthesamesensorypurposes。Immediatelyafterthemouthcomestheintestine,whichisneverwantinginanyinsect。Thisrunsinastraightlineandwithoutfurthercomplicationtothevent;

occasionally,however,ithasaspiralcoil。Thereare,moreover,someinsectsinwhichastomachsucceedstothemouth,andisitselfsucceededbyaconvolutedintestine,sothatthelargerandmorevoraciousinsectsmaybeenabledtotakeinamoreabundantsupplyoffood。MorecuriousthananyaretheCicadae。Forherethemouthandthetongueareunitedsoastoformasinglepart,throughwhich,asthrougharoot,theinsectsucksupthefluidsonwhichitlives。

Insectsarealwayssmalleaters,notsomuchbecauseoftheirdiminutivesizeasbecauseoftheircoldtemperament。Foritisheatwhichrequiressustenance;justasitisheatwhichspeedilyconcoctsit。Butcoldrequiresnosustenance。InnoinsectsisthissoconspicuousasintheseCicadae。Fortheyfindenoughtoliveoninthemoisturewhichisdepositedfromtheair。SoalsodotheEphemerathatarefoundabouttheBlacksea。Butwhiletheselatteronlyliveforasingleday,theCicadaesubsistonsuchfoodforseveraldays,thoughstillnotmany。

Wehavenowdonewiththeinternalpartsofanimals,andmustthereforereturntotheconsiderationoftheexternalpartswhichhavenotyetbeendescribed。Itwillbebettertochangeourorderofexpositionandbeginwiththeanimalswehavejustbeendescribing,sothatproceedingfromthese,whichrequirelessdiscussion,ouraccountmayhavemoretimetospendontheperfectkindsofanimals,thosenamelythathaveblood。

6

WewillbeginwithInsects。Theseanimals,thoughtheypresentnogreatmultiplicityofparts,arenotwithoutdiversitieswhencomparedwitheachother。Theyareallmanyfooted;theobjectofthisbeingtocompensatetheirnaturalslownessandfrigidity,andgivegreateractivitytotheirmotions。Accordinglywefindthatthosewhich,asthe(Millipedes),havelongbodies,andarethereforethemostliabletorefrigeration,havealsothegreatestnumberoffeet。Again,thebodyintheseanimalsisinsected-thereasonforthisbeingthattheyhavenotgotonevitalcentrebutmany-andthenumberoftheirfeetcorrespondstothatoftheinsections。

Shouldthefeetfallshortofthis,theirdeficiencyiscompensatedbythepowerofflight。Ofsuchflyinginsectssomeliveawanderinglife,andareforcedtomakelongexpeditionsinsearchoffood。Thesehaveabodyoflightweight,andfourfeathers,twooneitherside,tosupportit。Sucharebeesandtheinsectsakintothem。When,however,suchinsectsareofverysmallbulk,theirfeathersarereducedtotwo,asisthecasewithflies。Insectswithheavybodiesandofstationaryhabits,thoughnotpolypterousinthesamewayasbees,yethavesheathstotheirfeatherstomaintaintheirefficiency。SucharetheMelolonthaeandthelike。Fortheirstationaryhabitsexposetheirfeatherstomuchgreaterrisksthanarerunbythoseofinsectsthataremoreconstantlyinflight,andonthisaccounttheyareprovidedwiththisprotectingshield。Thefeatherofaninsecthasneitherbarbsnorshaft。For,thoughitiscalledafeather,itisnofeatheratall,butmerelyaskin-likemembranethat,owingtoitsdryness,necessarilybecomesdetachedfromthesurfaceofthebody,asthefleshysubstancegrowscold。

Theseanimalsthenhavetheirbodiesinsected,notonlyforthereasonsalreadyassigned,butalsotoenablethemtocurlroundinsuchamannerasmayprotectthemfrominjury;forsuchinsectsashavelongbodiescanrollthemselvesup,whichwouldbeimpossiblewereitnotfortheinsections;andthosethatcannotdothiscanyetdrawtheirsegmentsupintotheinsectedspaces,andsoincreasethehardnessoftheirbodies。Thiscanbefeltquiteplainlybyputtingthefingerononeoftheinsects,forinstance,knownasCanthari。Thetouchfrightenstheinsect,anditremainsmotionless,whileitsbodybecomeshard。Thedivisionofthebodyintosegmentsisalsoanecessaryresultoftherebeingseveralsupremeorgansinplaceofone;andthisagainisapartoftheessentialconstitutionofinsects,andisacharacterwhichapproximatesthemtoplants。Forasplants,thoughcutintopieces,canstilllive,soalsocaninsects。Thereis,however,thisdifferencebetweenthetwocases,thattheportionsofthedividedinsectliveonlyforalimitedtime,whereastheportionsoftheplantliveonandattaintheperfectformofthewhole,sothatfromonesingleplantyoumayobtaintwoormore。

Someinsectsarealsoprovidedwithanothermeansofprotectionagainsttheirenemies,namelyasting。Insomethisisinfront,connectedwiththetongue,inothersbehindattheposteriorend。

Forjustastheorganofsmellinelephantsanswersseveraluses,servingalikeasaweaponandforpurposesofnutrition,sodoesalsothesting,whenplacedinconnexionwiththetongue,asinsomeinsects,answermorethanoneend。Foritistheinstrumentthroughwhichtheyderivetheirsensationsoffood,aswellasthatwithwhichtheysuckitupandbringittothemouth。Suchoftheseinsectsashavenoanteriorstingareprovidedwithteeth,whichserveinsomeofthemforbitingthefood,andinothersforitsprehensionandconveyancetothemouth。Sucharetheiruses,forinstance,inantsandallthevariouskindsofbees。Asfortheinsectsthathaveastingbehind,thisweaponisgiventhembecausetheyareofafiercedisposition。Insomeofthemthestingislodgedinsidethebody,inbees,forexample,andwasps。Fortheseinsectsaremadeforflight,andweretheirstingexternalandofdelicatemakeitwouldsoongetspoiled;andif,ontheotherhand,itwereofthickerbuild,asinscorpions,itsweightwouldbeanincumbrance。Asforscorpionsthatliveonthegroundandhaveatail,theirstingmustbesetuponthis,asotherwiseitwouldbeofnouseasaweapon。Dipterousinsectsneverhaveaposteriorsting。Fortheveryreasonoftheirbeingdipterousisthattheyaresmallandweak,andthereforerequirenomorethantwofeatherstosupporttheirlightweight;andthesamereasonwhichreducestheirfeatherstotwocausestheirstingtobeinfront;fortheirstrengthisnotsufficienttoallowthemtostrikeefficientlywiththehinderpartofthebody。Polypterousinsects,ontheotherhand,areofgreaterbulk-indeeditisthiswhichcausesthemtohavesomanyfeathers;andtheirgreatersizemakesthemstrongerintheirhinderparts。Thestingofsuchinsectsisthereforeplacedbehind。Nowitisbetter,whenpossible,thatoneandthesameinstrumentshallnotbemadetoserveseveraldissimilaruses;butthatthereshallbeoneorgantoserveasaweapon,whichcanthenbeverysharp,andadistinctonetoserveasatongue,whichcanthenbeofspongytextureandfittoabsorbnutriment。Whenever,therefore,natureisabletoprovidetwoseparateinstrumentsfortwoseparateuses,withouttheonehamperingtheother,shedoesso,insteadofactinglikeacoppersmithwhoforcheapnessmakesaspitandlampholderinone。Itisonlywhenthisisimpossiblethatsheusesoneorganforseveralfunctions。

Theanteriorlegsareinsomecaseslongerthantheothers,thattheymayservetowipeawayanyforeignmatterthatmaylodgeontheinsect’seyesandobstructitssight,whichalreadyisnotverydistinctowingtotheeyesbeingmadeofahardsubstance。Fliesandbeesandthelikemaybeconstantlyseenthusdressingthemselveswithcrossedforelegs。Oftheotherlegs,thehinderarebiggerthanthemiddlepair,bothtoaidinrunningandalsothattheinsect,whenittakesflight,mayspringmoreeasilyfromtheground。Thisdifferenceisstillmoremarkedinsuchinsectsasleap,inlocustsforinstance,andinthevariouskindsoffleas。Forthesefirstbendandthenextendthelegs,and,bydoingso,arenecessarilyshotupfromtheground。Itisonlythe。hindlegsoflocusts,andnotthefrontones,thatresemblethesteeringoarsofaship。Forthisrequiresthatthejointshallbedeflectedinwards,andsuchisneverthecasewiththeanteriorlimbs。Thewholenumberoflegs,includingthoseusedinleaping,issixinalltheseinsects。

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IntheTestaceathebodyconsistsofbutfewparts,thereasonbeingthattheseanimalsliveastationarylife。Forsuchanimalsasmovemuchaboutmustofnecessityhavemorenumerouspartsthansuchasremainquiet;fortheiractivitiesaremany,andthemorediversifiedthemovementsthegreaterthenumberoforgansrequiredtoeffectthem。SomespeciesofTestaceaareabsolutelymotionless,andothersnotquitebutnearlyso。Nature,however,hasprovidedthemwithaprotectioninthehardnessoftheshellwithwhichshehasinvestedtheirbody。Thisshell,asalreadyhasbeensaid,mayhaveonevalve,ortwovalves,orbeturbinate。Inthelattercaseitmaybeeitherspiral,asinwhelks,ormerelyglobular,asinsea-urchins。

Whenithastwovalves,thesemaybegaping,asinscallopsandmussels,wherethevalvesareunitedtogetherononesideonly,soastoopenandshutontheother;ortheymaybeunitedtogetheronbothsides,asintheSolens(razor-fishes)。InallcasesaliketheTestaceahave,likeplants,theheaddownwards。Thereasonforthisis,thattheytakeintheirnourishmentfrombelow,justasdoplantswiththeirroots。Thustheunderpartscomeinthemtobeabove,andtheupperpartstobebelow。Thebodyisenclosedinamembrane,andthroughthistheanimalfiltersfluidfreefromsaltandabsorbsitsnutriment。Inallthereisahead;butnoneoftheparts,exceptingthisrecipientoffood,hasanydistinctivename。

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AlltheCrustaceacancrawlaswellasswim,andaccordinglytheyareprovidedwithnumerousfeet。Therearefourmaingenera,viz。

theCarabi,astheyarecalled,theAstaci,theCarides,andtheCarcini。Ineachofthesegenera,again,therearenumerousspecies,whichdifferfromeachothernotonlyasregardsshape,butalsoveryconsiderablyasregardssize。For,whileinsomespeciestheindividualsarelarge,inotherstheyareexcessivelyminute。TheCarcinoidandCaraboidCrustacearesembleeachotherinpossessingclaws。Theseclawsarenotforlocomotion,buttoserveinplaceofhandsforseizingandholdingobjects;andtheyarethereforebentintheoppositedirectiontothefeet,beingsotwistedastoturntheirconvexitytowardsthebody,whiletheirfeetturntowardsittheirconcavity。Forinthispositiontheclawsarebestsuitedforlayingholdofthefoodandcarryingittothemouth。ThedistinctionbetweentheCarabiandtheCarcini(Crabs)consistsintheformerhavingatailwhilethelatterhavenone。FortheCarabiswimaboutandatailisthereforeofusetothem,servingfortheirpropulsionlikethebladeofanoar。ButitwouldbeofnousetotheCrabs;fortheseanimalslivehabituallyclosetotheshore,andcreepintoholesandcorners。Insuchofthemasliveoutatsea,thefeetaremuchlessadaptedforlocomotionthanintherest,becausetheyarelittlegiventomovingaboutbutdependforprotectionontheirshell-likecovering。TheMaiaeandthecrabsknownasHeracleoticareexamplesofthis;thelegsintheformerbeingverythin,inthelatterveryshort。

Theveryminutecrabsthatarefoundamongthesmallfryatthebottomofthenethavetheirhindermostfeetflattenedoutintothesemblanceoffinsoroar-blades,soastohelptheanimalinswimming。

TheCaridesaredistinguishedfromtheCarcinoidspeciesbythepresenceofatail;andfromtheCaraboidsbytheabsenceofclaws。

Thisisexplainedbytheirlargenumberoffeet,onwhichhasbeenexpendedthematerialforthegrowthofclaws。Theirfeetagainarenumeroustosuittheirmodeofprogression,whichismainlybyswimming。

Ofthepartsontheventralsurface,thoseneartheheadareinsomeoftheseanimalsformedlikegills,fortheadmissionanddischargeofwater;whilethepartslowerdowndifferinthetwosexes。ForinthefemaleCarabithesearemorelaminarthaninthemales,andinthefemalecrabstheflapisfurnishedwithhairierappendages。Thisgivesamplerspaceforthedisposaloftheova,whichthefemalesretaininthesepartsinsteadoflettingthemgofree,asdofishesandallotheroviparousanimals。IntheCarabiandintheCrabstherightclawisinvariablythelargerandthestronger。Foritisnaturaltoeveryanimalinactiveoperationstousethepartsonitsrightsideinpreferencetothoseonitsleft;andnature,indistributingtheorgans,invariablyassignseach,eitherexclusivelyorinamoreperfectcondition,tosuchanimalsascanuseit。Soitiswithtusks,andteeth,andhorns,andspurs,andallsuchdefensiveandoffensiveweapons。

IntheLobstersaloneitisamatterofchancewhichclawisthelarger,andthisineithersex。Clawstheymusthave,becausetheybelongtoagenusinwhichthisisaconstantcharacter;buttheyhavetheminthisindeterminateway,owingtoimperfectformationandtotheirnotusingthemfortheirnaturalpurpose,butforlocomotion。

Foradetailedaccountoftheseveralpartsoftheseanimals,oftheirpositionandtheirdifferences,thosepartsbeingalsoincludedwhichdistinguishthesexes,referencemustbemadetothetreatisesonAnatomyandtotheResearchesconcerningAnimals。

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WecomenowtotheCephalopoda。Theirinternalorganshavealreadybeendescribedwiththoseofotheranimals。Externallythereisthetrunkofthebody,notdistinctlydefined,andinfrontofthistheheadsurroundedbyfeet,whichformacircleaboutthemouthandteeth,andaresetbetweentheseandtheeyes。Nowinallotheranimalsthefeet,ifthereareany,aredisposedinoneoftwoways;

eitherbeforeandbehindoralongthesides,thelatterbeingtheplaninsuchofthem,forinstance,asarebloodlessandhavenumerousfeet。ButintheCephalopodathereisapeculiararrangement,differentfromeitherofthese。Fortheirfeetareallplacedatwhatmaybecalledtheforeend。Thereasonforthisisthatthehindpartoftheirbodyhasbeendrawnupclosetotheforepart,asisalsothecaseintheturbinatedTestacea。FortheTestacea,whileinsomepointstheyresembletheCrustacea,inothersresembletheCephalopoda。Theirearthymatterisontheoutside,andtheirfleshysubstancewithin。SofartheyareliketheCrustacea。ButthegeneralplanoftheirbodyisthatoftheCephalopoda;and,thoughthisistrueinacertaindegreeofalltheTestacea,itismoreespeciallytrueofthoseturbinatedspeciesthathaveaspiralshell。Ofthisgeneralplan,commontothetwo,wewillspeakpresently。Butletusfirstconsiderthecaseofquadrupedsandofman,wherethearrangementisthatofastraightline。LetAattheupperendofsuchalinebesupposedtorepresentthemouth,thenB

thegullet,andCthestomach,andtheintestinetorunfromthisCtotheexcrementalventwhereDisinscribed。Suchistheplaninsanguineousanimals;androundthisstraightlineasanaxisaredisposedtheheadandso-calledtrunk;theremainingparts,suchastheanteriorandposteriorlimbs,havingbeensuperaddedbynature,merelytoministertotheseandforlocomotion。

IntheCrustaceaalsoandinInsectsthereisatendencytoasimilararrangementoftheinternalpartsinastraightline;thedistinctionbetweenthesegroupsandthesanguineousanimalsdependingondifferencesoftheexternalorganswhichministertolocomotion。

ButtheCephalopodaandtheturbinatedTestaceahaveincommonanarrangementwhichstandsincontrastwiththis。Forherethetwoextremitiesarebroughttogetherbyacurve,asifoneweretobendthestraightlinemarkedEuntilDcameclosetoSuch,then,isthedispositionoftheinternalparts;androundthese,intheCephalopoda,isplacedthesac(inthePoulpsalonecalledahead),and,intheTestacea,theturbinateshellwhichcorrespondstothesac。Thereis,infact,onlythisdifferencebetweenthem,thattheinvestingsubstanceoftheCephalopodaissoftwhiletheshelloftheTestaceaishard,naturehavingsurroundedtheirfleshypartwiththishardcoatingasaprotectionbecauseoftheirlimitedpoweroflocomotion。Inbothclasses,owingtothisarrangementoftheinternalorgans,theexcrementisvoidednearthemouth;atapointbelowthisorificeintheCephalopoda,andintheTurbinataononesideofit。

Such,then,istheexplanationofthepositionofthefeetintheCephalopoda,andofthecontrasttheypresenttootheranimalsinthismatter。Thearrangement,however,intheSepiasandtheCalamariesisnotpreciselythesameasinthePoulps,owingtotheformerhavingnoothermodeofprogressionthanbyswimming,whilethelatternotonlyswimbutcrawl。Forintheformersixofthefeetareabovetheteethandsmall,theouteroneoneithersidebeingthebiggest;

whiletheremainingtwo,whichmakeupthetotalweight,arebelowthemouthandarethebiggestofall,justasthehindlimbsinquadrupedsarestrongerthantheforelimbs。Foritisthesethathavetosupporttheweight,andtotakethemainpartinlocomotion。Andtheoutertwooftheuppersixarebiggerthanthepairwhichintervenebetweenthemandtheuppermostofall,becausetheyhavetoassistthelowermostpairintheiroffice。InthePoulps,ontheotherhand,thefourcentralfeetarethebiggest。Again,thoughthenumberoffeetisthesameinalltheCephalopoda,namelyeight,theirlengthvariesindifferentkinds,beingshortintheSepiasandtheCalamaries,butgreaterinthePoulps。Forintheselatterthetrunkofthebodyisofsmallbulk,whileintheformeritisofconsiderablesize;

andsointheonecasenaturehasusedthematerialssubtractedfromthebodytogivelengthtothefeet,whileintheothershehasactedinpreciselytheoppositeway,andhasgiventothegrowthofthebodywhatshehasfirsttakenfromthefeet。ThePoulps,then,owingtothelengthoftheirfeet,cannotonlyswimbutcrawl,whereasintheothergenerathefeetareuselessforthelattermodeofprogression,beingsmallwhilethebulkofthebodyisconsiderable。Theseshortfeetwouldnotenabletheirpossessorstoclingtotherocksandkeepthemselvesfrombeingtornoffbythewaveswhentheserunhighintimesofstorm;neitherwouldtheyservetolayholdofobjectsatallremoteandbringthemin;but,tosupplythesedefects,theanimalisfurnishedwithtwolongproboscises,bywhichitcanmooritselfandrideatanchorlikeashipinroughweather。Thesesameprocessesservealsotocatchpreyatadistanceandtobringittothemouth。TheyaresousedbyboththeSepiasandtheCalamaries。InthePoulpsthefeetarethemselvesabletoperformtheseoffices,andthereareconsequentlynoproboscises。Proboscisesandtwiningtentacles,withacetabulasetuponthem,actinthesamewayandhavethesamestructureasthoseplaitedinstrumentswhichwereusedbyphysiciansofoldtoreducedislocationsofthefingers。Likethesetheyaremadebytheinterlacingoftheirfibres,andtheyactbypullinguponpiecesoffleshandyieldingsubstances。Fortheplaitedfibresencircleanobjectinaslackenedcondition,andwhentheyareputonthestretchtheygraspandclingtightlytowhateveritmaybethatisincontactwiththeirinnersurface。Since,then,theCephalopodahavenootherinstrumentswithwhichtoconveyanythingtothemselvesfromwithout,thaneithertwiningtentacles,asinsomespecies,orproboscisesasinothers,theyareprovidedwiththesetoserveashandsforoffenceanddefenceandothernecessaryuses。

TheacetabulaaresetindoublelineinalltheCephalopodaexceptinginonekindofpoulp,wherethereisbutasinglerow。Thelengthandtheslimnesswhichispartofthenatureofthiskindofpoulpexplaintheexception。Foranarrowspacecannotpossiblyadmitofmorethanasinglerow。Thisexceptionalcharacter,then,belongstothem,notbecauseitisthemostadvantageousarrangement,butbecauseitisthenecessaryconsequenceoftheiressentialspecificconstitution。

Inalltheseanimalsthereisafin,encirclingthesac。InthePoulpsandtheSepiasthisfinisunbrokenandcontinuous,asisalsothecaseinthelargercalamariesknownasTeuthi。Butinthesmallerkind,calledTeuthides,thefinisnotonlybroaderthanintheSepiasandthePoulps,whereitisverynarrow,but,moreover,doesnotencircletheentiresac,butonlybeginsinthemiddleoftheside。Theuseofthisfinistoenabletheanimaltoswim,andalsotodirectitscourse。Itacts,thatis,liketherump-feathersinbirds,orthetail-fininfishes。InnoneisitsosmallorsoindistinctasinthePoulps。Forinthesethebodyisofsmallbulkandcanbesteeredbythefeetsufficientlywellwithoutotherassistance。

TheInsects,theCrustacea,theTestacea,andtheCephalopoda,havenowbeendealtwithinturn;andtheirpartshavebeendescribed,whetherinternalorexternal。

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Wemustnowgobacktotheanimalsthathaveblood,andconsidersuchoftheirparts,alreadyenumerated,aswerebeforepassedover。

Wewilltaketheviviparousanimalsfirst,and,wehavedonewiththese,willpassontotheoviparous,andtreatoftheminlikemanner。

Thepartsthatborderonthehead,andonwhatisknownastheneckandthroat,havealreadybeentakenintoconsideration。Allanimalsthathavebloodhaveahead;whereasinsomebloodlessanimals,suchascrabs,thepartwhichrepresentsaheadisnotclearlydefined。Astotheneck,itispresentinalltheVivipara,butonlyinsomeoftheOvipara;forwhilethosethathavealungalsohaveaneck,thosethatdonotinhaletheouterairhavenone。Theheadexistsmainlyforthesakeofthebrain。Foreveryanimalthathasbloodmustofnecessityhaveabrain;andmust,moreover,forreasonsalreadygiven,haveitplacedinanoppositeregiontotheheart。Buttheheadhasalsobeenchosenbynatureasthepartinwhichtosetsomeofthesenses;becauseitsbloodismixedinsuchsuitableproportionsastoensuretheirtranquillityandprecision,whileatthesametimeitcansupplythebrainwithsuchwarmthasitrequires。Thereisyetathirdconstituentsuperaddedtothehead,namelythepartwhichministerstotheingestionoffood。Thishasbeenplacedherebynature,becausesuchasituationaccordsbestwiththegeneralconfigurationofthebody。Forthestomachcouldnotpossiblybeplacedabovetheheart,seeingthatthisisthesovereignorgan;andifplacedbelow,asinfactitis,thenthemouthcouldnotpossiblybeplacedtherealso。Forthiswouldhavenecessitatedagreatincreaseinthelengthofthebody;andthestomach,moreover,wouldhavebeenremovedtoofarfromthesourceofmotionandofconcoction。

Thehead,then,existsforthesakeofthesethreeparts;whiletheneck,again,existsforthesakeofthewindpipe。Foritactsasadefencetothisandtotheoesophagus,encirclingthemandkeepingthemfrominjury。Inallotheranimalsthisneckisflexibleandcontainsseveralvertebrae;butinwolvesandlionsitcontainsonlyasinglebone。Fortheobjectofnaturewastogivetheseanimalsanorganwhichshouldbeserviceableinthewayofstrength,ratherthanonethatshouldbeusefulforanyoftheotherpurposestowhichnecksaresubservient。

Continuouswiththeheadandneckisthetrunkwiththeanteriorlimbs。Inmantheforelegsandforefeetarereplacedbyarmsandbywhatwecallhands。Forofallanimalsmanalonestandserect,inaccordancewithhisgodlikenatureandessence。Foritisthefunctionofthegod-liketothinkandtobewise;andnoeasytaskwerethisundertheburdenofaheavybody,pressingdownfromaboveandobstructingbyitsweightthemotionsoftheintellectandofthegeneralsense。When,moreover,theweightandcorporealsubstancebecomeexcessive,thebodymustofnecessityinclinetowardstheground。Insuchcasesthereforenature,inordertogivesupporttothebody,hasreplacedthearmsandhandsbyforefeet,andhasthusconvertedtheanimalintoaquadruped。For,aseveryanimalthatwalksmustofnecessityhavethetwohinderfeet,suchananimalbecomesaquadruped,itsbodyincliningdownwardsinfrontfromtheweightwhichitssoulcannotsustain。Forallanimals,manaloneexcepted,aredwarf-likeinform。Forthedwarf-likeisthatinwhichtheupperpartislarge,whilethatwhichbearstheweightandisusedinprogressionissmall。Thisupperpartiswhatwecallthetrunk,whichreachesfromthemouthtothevent。Inmanitisdulyproportionatetothepartbelow,anddiminishesmuchinitscomparativesizeasthemanattainstofullgrowth。Butinhisinfancythecontraryobtains,andtheupperpartsarelarge,whilethelowerpartissmall;sothattheinfantcanonlycrawl,andisunabletowalk;nay,atfirstcannotevencrawl,butremainswithoutmotion。Forallchildrenaredwarfsinshape,butceasetobesoastheybecomemen,fromthegrowthoftheirlowerpart;whereasinquadrupedsthereverseoccurs,theirlowerpartsbeinglargestinyouth,andadvanceofyearsbringingincreasedgrowthabove,thatisinthetrunk,whichextendsfromtherumptothehead。Thusitisthatcoltsarescarcely,ifatall,belowfull-grownhorsesinheight;andthatwhilestillyoungtheycantouchtheirheadswiththehindlegs,thoughthisisnolongerpossiblewhentheyareolder。Such,then,istheformofanimalsthathaveeitherasolidoraclovenhoof。Butsuchasarepolydactylousandwithouthorns,thoughtheytooareofdwarf-likeshape,aresoinalessdegree;andthereforethegreatergrowthofthelowerpartsascomparedwiththeupperisalsosmall,beingproportionatetothissmallerdeficiency。

Dwarf-likeagainistheraceofbirdsandfishes;andsoinfact,asalreadyhasbeensaid,iseveryanimalthathasblood。Thisisthereasonwhynootheranimalissointelligentasman。Forevenamongmenthemselvesifwecomparechildrenwithadults,orsuchadultsasareofdwarf-likeshapewithsuchasarenot,wefindthat,whateverothersuperioritytheformermaypossess,theyareatanyratedeficientascomparedwiththelatterinintelligence。Theexplanation,asalreadystated,isthattheirpsychicalprincipleiscorporeal,andmuchimpededinitsmotions。Letnowafurtherdecreaseoccurintheelevatingheat,andafurtherincreaseintheearthymatter,andtheanimalsbecomesmallerinbulk,andtheirfeetmorenumerous,untilatalaterstagetheybecomeapodous,andextendedfulllengthontheground。Then,byfurthersmallsuccessionsofchange,theycometohavetheirprincipalorganbelow;andatlasttheircephalicpartbecomesmotionlessanddestituteofsensation。

Thustheanimalbecomesaplant,thathasitsupperpartsdownwardsanditslowerpartsabove。Forinplantstherootsaretheequivalentsofmouthandhead,whiletheseedhasanoppositesignificance,foritisproducedaboveittheextremitiesofthetwigs。

Thereasonshavenowbeenstatedwhysomeanimalshavemanyfeet,someonlytwo,andothersnone;why,also,somelivingthingsareplantsandothersanimals;and,lastly,whymanaloneofallanimalsstandserect。Standingthuserect,manhasnoneedoflegsinfront,andintheirsteadhasbeenendowedbynaturewitharmsandhands。NowitistheopinionofAnaxagorasthatthepossessionofthesehandsisthecauseofmanbeingofallanimalsthemostintelligent。Butitismorerationaltosupposethathisendowmentwithhandsistheconsequenceratherthanthecauseofhissuperiorintelligence。Forthehandsareinstrumentsororgans,andtheinvariableplanofnatureindistributingtheorgansistogiveeachtosuchanimalascanmakeuseofit;natureactinginthismatterasanyprudentmanwoulddo。Foritisabetterplantotakeapersonwhoisalreadyaflute-playerandgivehimaflute,thantotakeonewhopossessesafluteandteachhimtheartofflute-playing。Fornatureaddsthatwhichislesstothatwhichisgreaterandmoreimportant,andnotthatwhichismorevaluableandgreatertothatwhichisless。

Seeingthenthatsuchisthebettercourse,andseeingalsothatofwhatispossiblenatureinvariablybringsaboutthebest,wemustconcludethatmandoesnotowehissuperiorintelligencetohishands,buthishandstohissuperiorintelligence。Forthemostintelligentofanimalsistheonewhowouldputthemostorganstouse;andthehandisnottobelookedonasoneorganbutasmany;foritis,asitwere,aninstrumentforfurtherinstruments。Thisinstrument,therefore,-thehand-ofallinstrumentsthemostvariouslyserviceable,hasbeengivenbynaturetoman,theanimalofallanimalsthemostcapableofacquiringthemostvariedhandicrafts。